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The Project Gutenberg eBook of A New Species of Frog (Genus
Tomodactylus) from Western México
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Title: A New Species of Frog (Genus Tomodactylus) from Western México
Author: Robert G. Webb
Release date: February 3, 2010 [eBook #31175]
Most recently updated: January 6, 2021
Language: English
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*** START OF THE PROJECT GUTENBERG EBOOK A NEW SPECIES
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University of Kansas Publications
Museum of Natural History
Tomodactylus) from Western México
This eBook is for the use of anyone anywhere in the United States and most
other parts of the world at no cost and with almost no restrictions
whatsoever. You may copy it, give it away or re-use it under the terms of
the Project Gutenberg License included with this eBook or online at
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have to check the laws of the country where you are located before using
this eBook.
Title: A New Species of Frog (Genus Tomodactylus) from Western México
Author: Robert G. Webb
Release date: February 3, 2010 [eBook #31175]
Most recently updated: January 6, 2021
Language: English
Other information and formats: www.gutenberg.org/ebooks/31175
Credits: Produced by Chris Curnow, Joseph Cooper and the Online
Distributed Proofreading Team at https://www.pgdp.net
*** START OF THE PROJECT GUTENBERG EBOOK A NEW SPECIES
OF FROG (GENUS TOMODACTYLUS) FROM WESTERN MÉXICO
***
University of Kansas Publications
Museum of Natural History
Page 4
Volume 15, No. 3, pp. 175-181, 1 fig.
March 7, 1962
A New Species of Frog (Genus
Tomodactylus)
from Western México
BY
ROBERT G. WEBB
University of Kansas
Lawrence
1962
University of Kansas Publications, Museum of Natural History
Editors: E. Raymond Hall, Chairman, Henry S. Fitch,
Theodore H. Eaton, Jr.
Volume 15, No. 3, pp. 175-181, 1 fig.
Published March 7, 1962
University of Kansas
Lawrence, Kansas
March 7, 1962
A New Species of Frog (Genus
Tomodactylus)
from Western México
BY
ROBERT G. WEBB
University of Kansas
Lawrence
1962
University of Kansas Publications, Museum of Natural History
Editors: E. Raymond Hall, Chairman, Henry S. Fitch,
Theodore H. Eaton, Jr.
Volume 15, No. 3, pp. 175-181, 1 fig.
Published March 7, 1962
University of Kansas
Lawrence, Kansas
Page 5
PRINTED BY
JEAN M. NEIBARGER, STATE PRINTER
TOPEKA, KANSAS
1962
29-1923
JEAN M. NEIBARGER, STATE PRINTER
TOPEKA, KANSAS
1962
29-1923
Page 6
A New Species of Frog (Genus Tomodactylus)
from Western México.
BY
ROBERT G. WEBB
Thirteen specimens of frogs collected in the summers of 1960 and 1961 in
the Mexican states of Durango and Sinaloa represent a heretofore unnamed
species. The specimens have been deposited in the Museum of Natural
History of the University of Kansas (KU) and in the Museum of Michigan
State University (MSU). The species may be named and described as
follows:
Tomodactylus saxatilis new species
Holotype.—KU 63326 (Fig. 1); obtained eight miles west of El Palmito, Sinaloa, approximately 6100
feet, on 23 June 1961; original field number, 2354 of Robert G. Webb.
Paratypes.—A total of 12 specimens: KU 63327-33, same data as holotype, 23-25 June 1961; MSU
4085-88, two miles north of Pueblo Nuevo, Durango, approximately 6000 feet elevation, 24 July
1960; MSU 4089, one half mile west of Revolcaderos, Durango, approximately 6600 feet, 29 July
1961.
Diagnosis.—A species of Tomodactylus possessing the following combination of characters: (1) tips
of two outer fingers truncate, about twice width of narrowest part of digit; (2) tympanum small, less
than one half diameter of eye; (3) ventral surfaces smooth; (4) contrasting marbled pattern on back
and top of head, and (5) venter whitish, lacking dark marks.
Description of holotype.—Adult male; snout-vent length, 31.5 (measurements are in millimeters and
were taken by means of dial calipers reading to one tenth of a millimeter); width of head, 11.2; length
of head, 10.3; horizontal diameter of eye, 3.1, and of tympanum, 1.2; distance from eye to nostril,
3.8; internarial width, 2.9; interorbital width, 4.1; width of eyelid, 2.5; lumbar gland (left side), 7.0 x
2.6; distance from axilla to groin, 15.2; tibial length, 12.7; length of foot, 13.1.
Head slightly wider than body; tip of snout rounded, slightly truncate; canthus rounded; tympanum
small, less than one half diameter of eye; tympanum having posterior margin ill-defined, separated
from eye by distance about equal to diameter of tympanum; diameter of eye slightly less than
distance from eye to nostril; width of eyelid about two thirds interorbital width; paratoid gland
indistinct; lumbar glands high, separated from insertion of leg by about one millimeter; back and
sides of body having low, scarcely elevated pustules; top of head, limbs and venter smooth; few low,
whitish pustules below and behind tympanum, and low on sides of body; posterior surface of thighs
from Western México.
BY
ROBERT G. WEBB
Thirteen specimens of frogs collected in the summers of 1960 and 1961 in
the Mexican states of Durango and Sinaloa represent a heretofore unnamed
species. The specimens have been deposited in the Museum of Natural
History of the University of Kansas (KU) and in the Museum of Michigan
State University (MSU). The species may be named and described as
follows:
Tomodactylus saxatilis new species
Holotype.—KU 63326 (Fig. 1); obtained eight miles west of El Palmito, Sinaloa, approximately 6100
feet, on 23 June 1961; original field number, 2354 of Robert G. Webb.
Paratypes.—A total of 12 specimens: KU 63327-33, same data as holotype, 23-25 June 1961; MSU
4085-88, two miles north of Pueblo Nuevo, Durango, approximately 6000 feet elevation, 24 July
1960; MSU 4089, one half mile west of Revolcaderos, Durango, approximately 6600 feet, 29 July
1961.
Diagnosis.—A species of Tomodactylus possessing the following combination of characters: (1) tips
of two outer fingers truncate, about twice width of narrowest part of digit; (2) tympanum small, less
than one half diameter of eye; (3) ventral surfaces smooth; (4) contrasting marbled pattern on back
and top of head, and (5) venter whitish, lacking dark marks.
Description of holotype.—Adult male; snout-vent length, 31.5 (measurements are in millimeters and
were taken by means of dial calipers reading to one tenth of a millimeter); width of head, 11.2; length
of head, 10.3; horizontal diameter of eye, 3.1, and of tympanum, 1.2; distance from eye to nostril,
3.8; internarial width, 2.9; interorbital width, 4.1; width of eyelid, 2.5; lumbar gland (left side), 7.0 x
2.6; distance from axilla to groin, 15.2; tibial length, 12.7; length of foot, 13.1.
Head slightly wider than body; tip of snout rounded, slightly truncate; canthus rounded; tympanum
small, less than one half diameter of eye; tympanum having posterior margin ill-defined, separated
from eye by distance about equal to diameter of tympanum; diameter of eye slightly less than
distance from eye to nostril; width of eyelid about two thirds interorbital width; paratoid gland
indistinct; lumbar glands high, separated from insertion of leg by about one millimeter; back and
sides of body having low, scarcely elevated pustules; top of head, limbs and venter smooth; few low,
whitish pustules below and behind tympanum, and low on sides of body; posterior surface of thighs
Page 7
and anal region pustulate; one pair of whitish postanal spots; ventral disc attached near insertion of
legs, lacking conspicuous transverse fold; skin loose on throat, chest and abdomen.
Digits not webbed; tips of two outer fingers truncate, having terminal transverse grooves, about twice
width of narrowest part of digit; digits of first and second fingers slightly expanded; fingers from
shortest to longest, 1-2-4-3, first only slightly shorter than second; three palmar tubercles; inner
palmar tubercle about one third size of large median tubercle; outer tubercle about one tenth size of
large median tubercle; four supernumerary palmar tubercles; tips of toes slightly wider than
narrowest part of digits; toes from shortest to longest, 1-2-5-3-4, second only slightly shorter than
fifth; inner metatarsal tubercle about four times size of small outer metatarsal tubercle;
supernumerary tubercles on foot small; no tarsal fold; heels touching when tibiae adpressed to thighs;
tibiotarsal articulation reaching eye when leg adpressed to side of body.
Contrasting marbled pattern on back and top of head; contrasting, mostly barred, pattern on limbs;
ventral surfaces whitish, lacking dark marks, but having minute dark peppering; marbling of dorsal
surfaces blackish and whitish in preservative.
Vomerine teeth lacking; internal choanae lateral, partly concealed by maxillaries; tongue smooth,
elongate, shallowly notched distally, free for about half its length; vocal sac median; internal vocal
slits large and near angle of jaw.
Variation.—Twelve males closely resemble the holotype. Two specimens from Pueblo Nuevo are soft
and not well preserved. The ranges of variation (means in parentheses) for the 13 males comprising
the type series are: snout-vent length, 25.5-31.5 (27.9); width of head, 9.7-11.2 (10.5); diameter of
eye, 2.9-3.6 (3.2); horizontal diameter of tympanum, 1.1-1.8 (1.4); length of tibia, 11.1-13.1 (12.1);
length of foot, 11.0-13.1 (12.1).
The pale ground color of the marbled pattern in most specimens is least extensive on the back and
arms, but most extensive on the legs. The lumbar glands are slightly elevated and conspicuous, and in
KU 63328 are extremely protuberant, or (KU 63330) evident on left side but flattened and indistinct
on right side. The back is rough having low, scarcely elevated pustules, but becomes less rough
anteriorly and most of the top of head is smooth. The three specimens from Pueblo Nuevo, Durango,
differ slightly from the other specimens examined in lacking pairs of postanal white spots, and in
having smooth backs (slightly pustulate in MSU 4088). The tibiotarsal articulation fails to reach the
eye in KU 63330. The small inner palmar tubercle is continuous with the large median tubercle on
the right hand of KU 63330, and lacking on both hands of KU 63329 and on the left hand of KU
63328. The tip of the tongue is entire in some specimens and in others has an irregular margin.
Coloration of living specimens.—Marbled pattern on back and top of head of dark brown to blackish
on yellowish-gold; pattern slightly less contrasting on limbs than on back, consisting of brown to
grayish on pale yellow; side of head and body grayish sometimes having pale yellow to whitish
spots; iris blackish having fine reticulation of yellowish to greenish-gold; venter dirty white.
Habitat.—The three records of occurrence for Tomodactylus saxatilis are in
a mixed boreal-tropical habitat, which is transitional between a pine-oak
forest at higher elevations and a tropical deciduous forest at lower
elevations. The mixed boreal-tropical habitat is most conspicuous at
elevations between approximately 7800 and 5500 feet on southerly exposed
slopes of barrancas and arroyos of the dissected plateau of the Sierra Madre
Occidental. The mixed boreal-tropical habitat occurs for approximately 30
legs, lacking conspicuous transverse fold; skin loose on throat, chest and abdomen.
Digits not webbed; tips of two outer fingers truncate, having terminal transverse grooves, about twice
width of narrowest part of digit; digits of first and second fingers slightly expanded; fingers from
shortest to longest, 1-2-4-3, first only slightly shorter than second; three palmar tubercles; inner
palmar tubercle about one third size of large median tubercle; outer tubercle about one tenth size of
large median tubercle; four supernumerary palmar tubercles; tips of toes slightly wider than
narrowest part of digits; toes from shortest to longest, 1-2-5-3-4, second only slightly shorter than
fifth; inner metatarsal tubercle about four times size of small outer metatarsal tubercle;
supernumerary tubercles on foot small; no tarsal fold; heels touching when tibiae adpressed to thighs;
tibiotarsal articulation reaching eye when leg adpressed to side of body.
Contrasting marbled pattern on back and top of head; contrasting, mostly barred, pattern on limbs;
ventral surfaces whitish, lacking dark marks, but having minute dark peppering; marbling of dorsal
surfaces blackish and whitish in preservative.
Vomerine teeth lacking; internal choanae lateral, partly concealed by maxillaries; tongue smooth,
elongate, shallowly notched distally, free for about half its length; vocal sac median; internal vocal
slits large and near angle of jaw.
Variation.—Twelve males closely resemble the holotype. Two specimens from Pueblo Nuevo are soft
and not well preserved. The ranges of variation (means in parentheses) for the 13 males comprising
the type series are: snout-vent length, 25.5-31.5 (27.9); width of head, 9.7-11.2 (10.5); diameter of
eye, 2.9-3.6 (3.2); horizontal diameter of tympanum, 1.1-1.8 (1.4); length of tibia, 11.1-13.1 (12.1);
length of foot, 11.0-13.1 (12.1).
The pale ground color of the marbled pattern in most specimens is least extensive on the back and
arms, but most extensive on the legs. The lumbar glands are slightly elevated and conspicuous, and in
KU 63328 are extremely protuberant, or (KU 63330) evident on left side but flattened and indistinct
on right side. The back is rough having low, scarcely elevated pustules, but becomes less rough
anteriorly and most of the top of head is smooth. The three specimens from Pueblo Nuevo, Durango,
differ slightly from the other specimens examined in lacking pairs of postanal white spots, and in
having smooth backs (slightly pustulate in MSU 4088). The tibiotarsal articulation fails to reach the
eye in KU 63330. The small inner palmar tubercle is continuous with the large median tubercle on
the right hand of KU 63330, and lacking on both hands of KU 63329 and on the left hand of KU
63328. The tip of the tongue is entire in some specimens and in others has an irregular margin.
Coloration of living specimens.—Marbled pattern on back and top of head of dark brown to blackish
on yellowish-gold; pattern slightly less contrasting on limbs than on back, consisting of brown to
grayish on pale yellow; side of head and body grayish sometimes having pale yellow to whitish
spots; iris blackish having fine reticulation of yellowish to greenish-gold; venter dirty white.
Habitat.—The three records of occurrence for Tomodactylus saxatilis are in
a mixed boreal-tropical habitat, which is transitional between a pine-oak
forest at higher elevations and a tropical deciduous forest at lower
elevations. The mixed boreal-tropical habitat is most conspicuous at
elevations between approximately 7800 and 5500 feet on southerly exposed
slopes of barrancas and arroyos of the dissected plateau of the Sierra Madre
Occidental. The mixed boreal-tropical habitat occurs for approximately 30
Page 8
miles along the paved highway (Mexican Highway 40) between Cd.
Durango, Durango, and Mazatlán, Sinaloa. The records of occurrence in
those states that are along this highway are separated by 14.5 miles (via
road).
Fig. 1. Tomodactylus saxatilis new species,
adult male, KU 63326, holotype (× 2),
dorsal view.
The terrain consists of occasional level areas, but is mostly of steep
hillsides. Dominant trees are large oaks and pines; a characteristic pine is
the sad or drooping-needle pine, locally called "pino triste." The
vegetational cover is usually open, including grasses, small oaks and pines,
broad-leaved shrubs and herbs, prickly pears, magueys, thorny acacias,
bracken fern, and epiphytes in trees. Ferns occur in moist protected places,
and orchids are occasional, sometimes in trees.
Outcrops of rock, boulder-strewn areas, and occasional rock slides (talus)
also characterize the terrain. Tomodactylus saxatilis seems to be restricted
to rocky habitats. The individuals collected were detected when they called
at night from within crevices of rocks or from exposed perches on rocks and
boulders; some calling frogs were out of reach on steep rock walls. The call
is a single, loud, high peep.
Durango, Durango, and Mazatlán, Sinaloa. The records of occurrence in
those states that are along this highway are separated by 14.5 miles (via
road).
Fig. 1. Tomodactylus saxatilis new species,
adult male, KU 63326, holotype (× 2),
dorsal view.
The terrain consists of occasional level areas, but is mostly of steep
hillsides. Dominant trees are large oaks and pines; a characteristic pine is
the sad or drooping-needle pine, locally called "pino triste." The
vegetational cover is usually open, including grasses, small oaks and pines,
broad-leaved shrubs and herbs, prickly pears, magueys, thorny acacias,
bracken fern, and epiphytes in trees. Ferns occur in moist protected places,
and orchids are occasional, sometimes in trees.
Outcrops of rock, boulder-strewn areas, and occasional rock slides (talus)
also characterize the terrain. Tomodactylus saxatilis seems to be restricted
to rocky habitats. The individuals collected were detected when they called
at night from within crevices of rocks or from exposed perches on rocks and
boulders; some calling frogs were out of reach on steep rock walls. The call
is a single, loud, high peep.
Page 9
Comparisons.—Dixon (1957) recognized six species of Tomodactylus (nitidus, dilatus, albolabris,
angustidigitorum, fuscus and grandis) in his revision of the genus. Another species (rufescens) was
subsequently described by Duellman and Dixon (1959). Tomodactylus saxatilis differs from all the
species named immediately above by the combination of characters given in the diagnosis.
Tomodactylus saxatilis differs from nitidus, angustidigitorum and grandis in having the tips of the
two outer fingers widened and truncate; saxatilis differs from dilatus, albolabris, fuscus and
rufescens in having a smooth venter (not pustulate), a contrasting pale and dark marbled pattern on
the back, and a lack of "flash" colors on the femora.
Tomodactylus saxatilis, having lumbar glands, also resembles three species referred to the genus
Syrrhophus. Tomodactylus macrotympanum was described by Taylor (1940:496, 497) as having a
large, moderately distinct lumbar gland; the species was referred to the genus Syrrhophus by Dixon
(op. cit.:384). According to Firschein (1954:55), Syrrhophus smithi and S. petrophilus have elongate
lumbar glands shaped like those in Tomodactylus. Tomodactylus saxatilis resembles macrotympanum,
smithi and petrophilus more than it does other species; all four attain large maximal sizes, and have
lumbar glands, mostly smooth ventral surfaces, three palmar tubercles (sometimes absent in
saxatilis), and usually contrasting dorsal patterns (reduced to flecks and spots in all species except
saxatilis). Tomodactylus saxatilis differs from macrotympanum in having an extensive marbled dorsal
pattern and a small tympanum, and differs from smithi and petrophilus in having a marbled dorsal
pattern and the tips of the outer two fingers widened and truncate. Tomodactylus saxatilis differs
from all other named species of Syrrhophus in having conspicuous lumbar glands and in lacking
inguinal glands.
Remarks.—The characteristics delimiting the genera Tomodactylus and
Syrrhophus are not agreed upon by all workers (see discussions by
Firschein, 1954:50; Langebartel and Shannon, 1956:164; and Dixon,
1957:383). I have referred saxatilis to the genus Tomodactylus on the basis
of a lumbar gland, which was considered a distinguishing character for the
genus by Smith and Taylor (1948:46) and Langebartel and Shannon
(1956:165). Lumbar glands are longer than broad, at least one third the
distance from axilla to groin, lateral and usually high, and often
conspicuous and protuberant. The elevation or flatness of the lumbar glands
seems to be due to individual variation; living specimens in the field had
conspicuous and protuberant, or non-elevated, indistinct lumbar glands.
Lumbar glands are not to be confused with inguinal glands, which are
roundish, often yellowish, sometimes diffuse, lateral but low, often
inconspicuous, and usually not protuberant. Inguinal glands occur in the
genus Microbatrachylus and in some species of Eleutherodactylus, and
have been described as flat, or low, or small, or indistinct for most species
of Syrrhophus.
For financial assistance with field work I am grateful to Rollin H. Baker,
and those individuals who administer the Michigan State University
Development Fund and the Bache Fund of the National Academy of
angustidigitorum, fuscus and grandis) in his revision of the genus. Another species (rufescens) was
subsequently described by Duellman and Dixon (1959). Tomodactylus saxatilis differs from all the
species named immediately above by the combination of characters given in the diagnosis.
Tomodactylus saxatilis differs from nitidus, angustidigitorum and grandis in having the tips of the
two outer fingers widened and truncate; saxatilis differs from dilatus, albolabris, fuscus and
rufescens in having a smooth venter (not pustulate), a contrasting pale and dark marbled pattern on
the back, and a lack of "flash" colors on the femora.
Tomodactylus saxatilis, having lumbar glands, also resembles three species referred to the genus
Syrrhophus. Tomodactylus macrotympanum was described by Taylor (1940:496, 497) as having a
large, moderately distinct lumbar gland; the species was referred to the genus Syrrhophus by Dixon
(op. cit.:384). According to Firschein (1954:55), Syrrhophus smithi and S. petrophilus have elongate
lumbar glands shaped like those in Tomodactylus. Tomodactylus saxatilis resembles macrotympanum,
smithi and petrophilus more than it does other species; all four attain large maximal sizes, and have
lumbar glands, mostly smooth ventral surfaces, three palmar tubercles (sometimes absent in
saxatilis), and usually contrasting dorsal patterns (reduced to flecks and spots in all species except
saxatilis). Tomodactylus saxatilis differs from macrotympanum in having an extensive marbled dorsal
pattern and a small tympanum, and differs from smithi and petrophilus in having a marbled dorsal
pattern and the tips of the outer two fingers widened and truncate. Tomodactylus saxatilis differs
from all other named species of Syrrhophus in having conspicuous lumbar glands and in lacking
inguinal glands.
Remarks.—The characteristics delimiting the genera Tomodactylus and
Syrrhophus are not agreed upon by all workers (see discussions by
Firschein, 1954:50; Langebartel and Shannon, 1956:164; and Dixon,
1957:383). I have referred saxatilis to the genus Tomodactylus on the basis
of a lumbar gland, which was considered a distinguishing character for the
genus by Smith and Taylor (1948:46) and Langebartel and Shannon
(1956:165). Lumbar glands are longer than broad, at least one third the
distance from axilla to groin, lateral and usually high, and often
conspicuous and protuberant. The elevation or flatness of the lumbar glands
seems to be due to individual variation; living specimens in the field had
conspicuous and protuberant, or non-elevated, indistinct lumbar glands.
Lumbar glands are not to be confused with inguinal glands, which are
roundish, often yellowish, sometimes diffuse, lateral but low, often
inconspicuous, and usually not protuberant. Inguinal glands occur in the
genus Microbatrachylus and in some species of Eleutherodactylus, and
have been described as flat, or low, or small, or indistinct for most species
of Syrrhophus.
For financial assistance with field work I am grateful to Rollin H. Baker,
and those individuals who administer the Michigan State University
Development Fund and the Bache Fund of the National Academy of
Page 10
Sciences (Grant No. 463). I am grateful also to J. Keever Greer, Donald F.
Switzenberg, and Rudolph A. Scheibner for aid in the field, to Edward H.
Taylor, James R. Dixon, and William E. Duellman for profitable
discussions, and to Thomas Sweringen for figure 1. The specific name
alludes to the habitat (Latin, saxatilis = found among rocks).
LITERATURE CITED
Dixon, J. R.
1957. Geographic variation and distribution of the genus Tomodactylus in Mexico.
Texas Journ. Sci., 9(4):379-409, 5 figs., 1 map, December.
Duellman, W. E., and Dixon, J. R.
1959. A new frog of the genus Tomodactylus from Michoacan, Mexico. Texas Journ.
Sci., 11(1):78-82, 1 fig., 1 table, March.
Firschein, I. L.
1954. Definition of some little-understood members of the leptodactylid genus
Syrrhophus, with a description of a new species. Copeia, 1954, 1:48-58,
February 19.
Langebartel, D. A., and Shannon, F. A.
1956. A new frog (Syrrhophus) from the Sinaloan lowlands of Mexico. Herpetologica,
12(3):163-165, 2 figs., September 1.
Smith, H. M., and Taylor, E. H.
1948. An annotated checklist and key to the Amphibia of Mexico. Bull. U. S. Nat.
Mus., No. 194, pp. iv + 118.
Taylor, E. H.
1940. Herpetological miscellany. Univ. Kansas Sci. Bull., 26(15):489-571, 10 pls., 7
figs., November 15.
29-1923
Switzenberg, and Rudolph A. Scheibner for aid in the field, to Edward H.
Taylor, James R. Dixon, and William E. Duellman for profitable
discussions, and to Thomas Sweringen for figure 1. The specific name
alludes to the habitat (Latin, saxatilis = found among rocks).
LITERATURE CITED
Dixon, J. R.
1957. Geographic variation and distribution of the genus Tomodactylus in Mexico.
Texas Journ. Sci., 9(4):379-409, 5 figs., 1 map, December.
Duellman, W. E., and Dixon, J. R.
1959. A new frog of the genus Tomodactylus from Michoacan, Mexico. Texas Journ.
Sci., 11(1):78-82, 1 fig., 1 table, March.
Firschein, I. L.
1954. Definition of some little-understood members of the leptodactylid genus
Syrrhophus, with a description of a new species. Copeia, 1954, 1:48-58,
February 19.
Langebartel, D. A., and Shannon, F. A.
1956. A new frog (Syrrhophus) from the Sinaloan lowlands of Mexico. Herpetologica,
12(3):163-165, 2 figs., September 1.
Smith, H. M., and Taylor, E. H.
1948. An annotated checklist and key to the Amphibia of Mexico. Bull. U. S. Nat.
Mus., No. 194, pp. iv + 118.
Taylor, E. H.
1940. Herpetological miscellany. Univ. Kansas Sci. Bull., 26(15):489-571, 10 pls., 7
figs., November 15.
29-1923
Page 11
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OF FROG (GENUS TOMODACTYLUS) FROM WESTERN MÉXICO
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